Katsuki, T., Greenspan, R.J. (2013) Jellyfish nervous systems. Current Biology 23: R592–R594.

Greenspan, R.J.  (2009)  The landscape of selection, gene interaction, and gene networks. Cold Spring Harbor Symp. Quant. Biol. 74: 131-138.

Foltenyi, K., Greenspan, R.J., Newport, J.W. (2007) Activation of Egf-r and ERK by Rhomboid signaling regulates the consolidation and maintenance of sleep in Drosophila. Nature Neuroscience  10: 1160 – 1167.

Dierick, H.A., Greenspan, R.J. (2007) Serotonin and neuropeptide F have opposite modulatory effects on fly aggression. Nature Genetics 39: 678-682.

Dierick, H.A., Greenspan, R.J. (2006) Molecular analysis of flies selected for aggressive behavior.    Nature Genetics 38: 1023-1031.

Kendler, K.S., Greenspan, R.J. (2006) The nature of genetic influences on behavior: Lessons from “simpler” organisms.  Am. J. Psychiatry 163: 1683-1694.

Andretic, R., van Swinderen, B., Greenspan, R.J. (2005) Dopaminergic modulation of arousal in Drosophila.  Current Biology 15: 1165-1175.

Greenspan, R.J., van Swinderen, B. (2004)  Cognitive consonance: Complex brain functions in the fruit fly and its relatives.  Trends in Neurosciences 27: 707-711.

van Swinderen, B., Greenspan, R.J.  (2003) Salience Modulates 20-30 Hz Brain Activity in Drosophila. Nature Neuroscience 6: 579-586.

Toma, D.P., White, K.P., Hirsch, J., Greenspan, R.J.  (2002)  Identification of genes involved in Drosophila melanogaster geotaxis, a complex behavioral trait.  Nature Genetics 31: 349-353.

Greenspan, R.J. (2001)  The flexible genome.  Nature Reviews Genetics 2: 383-387.

Shaw, P.J., Cirelli, C., Greenspan, R.J., Tononi, G. (2000) Correlates of sleep and waking in Drosophila melanogaster.  Science 287: 1834-1837.

Osborne, K.A., Robichon, A., Burgess, E., Butland, S.,  Shaw, R.A., Coulthard, A., Pererira, Greenspan, R.J., Sokolowski, M.B. (1997) Natural behavior polymorphism due to a cGMP-dependent protein kinase of Drosophila.  Science 277: 834-836.

Ferveur, J.-F., Störtkuhl, K., Stocker, R.F., Greenspan, R.J.  (1995) Genetic feminization of brain structures and changed sexual orientation in male Drosophila melanogaster.   Science  267: 902-905.

Greenspan, R.J. (1995)  Understanding the genetic construction of behavior.  Scientific American  272 (4): 72-78.


Flint, J., Greenspan, R.J., and Kendler, K.S. (2010) How Genes Influence Behavior.  Oxford U. Press: London & New York.

Greenspan, R.J. (2007)  An Introduction to Nervous Systems.  Cold Spring Harbor Laboratory Press: Cold Spring Harbor, NY.